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Ecology and Conservation

Professor Richard (Dez) Delahay

Professor Richard (Dez) Delahay

Honorary Visiting Professor

 R.Delahay@exeter.ac.uk

 Environment and Sustainability Institute 

 

Environment and Sustainability Institute, University of Exeter, Penryn Campus, Penryn, Cornwall, TR10 9FE, UK


Overview

I am a wildlife biologist working at the interface between ecology and epidemiology. In my role as a scientist for the Animal and Plant Health Agency I work mainly on the ecology and management of diseases in wildlife, particularly bovine tuberculosis (TB). I manage research and oversee the fieldwork associated with a long-term capture-mark-recapture study of badgers at Woodchester Park, Gloucestershire. The project is a strategically important resource and has provided many insights into the epidemiology and management of TB in badgers and cattle. 

In my role as an APHA scientist I am involved in providing advice to Government and other stakeholders. I work closely with academic colleagues in the Centre for Ecology & Conservation and the Environment and Sustainability Institute

Research supervision

I co-supervise PhD researchers working on a variety of topics related to mammal ecology, and the epidemiology and management of wildlife diseases. Recent projects have investigated the spatio-temporal dynamics of TB transmission, senescence and immuno-competence and genetic correlates of TB infection in badgers, contact chains and disease risks in cattle, and the implications of contact networks for disease management in wild geese.  

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Research

Research interests

My research interests include mammal ecology and the epidemiology and management of wildlife diseases.

Publications

Reports/proceedings

Cromie, R.L., Lee, R., Delahay, R.J., Newth, J.L, O’Brien, M.F., Fairlamb, H.A., Reeves, J.P. & Stroud, D.A. (2012). Ramsar Wetland Disease Manual: Guidelines for Assessment, Monitoring and Management of Animal Disease in Wetlands. Ramsar Technical Report No.7. Ramsar Convention Secretariat, Gland, Switzerland.

Delahay, R. J. & Spray, C. J. (Eds) (2015).  Proceedings of the Oxford Lead Symposium. Lead Ammunition: understanding and minimising the risks to human and environmental health.  Edward Grey Institute, The University of Oxford, UK.  152pp.

Books (Ed.)

Delahay, R. J., Smith, G. C. & Hutchings, M. R. (2008).  Management of Disease in Wild Mammals. Springer Tokyo.  pp284.

Book chapters

Rogers, L. M., Delahay, R. J., Hounsome, T. D. & Cheeseman, C. L. (2000).  Changes in badger (Meles meles) social organisation in response to increasing population density at Woodchester Park, south west England.  In Mustelids in a Modern World: Management and conservation aspects of small carnivore:human interactions. Ed. H. I. Griffiths.  Backhuys Publishers, Leiden, Nederlands.   pp342.

Delahay, R. J., Wilson, G., Rogers, L. M. & Cheeseman, C. L. (2003). TB and the badger: Culling as a method to control a wildlife disease reservoir. In Conservation and Conflict: Mammals and Farming in Britain. F.H. Tattersall & W.J. Manley (eds) Linnean Society Occasional Publication Series, Smith Settle: Otley.  p165-171.

Delahay, R. J., Wilson, G., Harris, S. & Macdonald, D.  (2008).  Badger Meles meles. In Mammals of the British Isles Handbook, 4th Edition. S. Harris & D.W. Yalden (eds) The Mammal Society. p425-436.

Delahay, R. J., Smith, G. C. & Hutchings, M. R. (2009).  The science of wildlife disease management.  In Management of Disease in Wild Mammals. R. J. Delahay, G. C. Smith & M. R. Hutchings (eds). Springer Tokyo.  p1-8.

Cross, P. C., Drewe, J., Patrek, V., Pearce, G., Samuel, M. D. & Delahay, R. J. (2009).  Wildlife population structure and parasite transmission: implications for disease management.  In Management of Disease in Wild Mammals. R. J. Delahay, G. C. Smith & M. R. Hutchings (eds). Springer Tokyo.  p9-29.

Blancou, J., Artois, M., Gilot-Fromont, E., Kaden, V., Rossi, S., Smith, G. C., Hutchings, M. R., Chambers, M. A., Houghton, S. & Delahay, R. J. (2009).  Options for the control of disease: targeting the infectious or parasitic agent.  In Management of Disease in Wild Mammals. R. J. Delahay, G. C. Smith & M. R. Hutchings (eds). Springer Tokyo.  p97-120.

Carter, S. P., Roy, S. S., Cowan, D. P., Massei, G., Smith, G. C., Ji, W., Rossi, S., Woodroffe, R., Wilson, G. J. & Delahay, R. J.  (2009).  Options for the control of disease 2: targeting hosts.  In Management of Disease in Wild Mammals. R. J. Delahay, G. C. Smith & M. R. Hutchings (eds). Springer Tokyo.  p121-146.

Ward, A. I., VerCauteren, K. C., Walter, W. D., Gilot-Fromont, E., Rossi, S., Edwards-Jones, G., Lambert, M., Hutchings, M. R. & Delahay, R. J.  (2009).  Options for the control of disease 3: targeting the environment.  In Management of Disease in Wild Mammals. R. J. Delahay, G. C. Smith & M. R. Hutchings (eds). Springer Tokyo.  p147-168.

Jackson, V. S., Huntley, S., Tomlinson, A., Smith, G. C., Taylor, M. A. & Delahay, R. J.  (2009).  Risk assessment and contingency planning for exotic disease introductions.  In Management of Disease in Wild Mammals. R. J. Delahay, G. C. Smith & M. R. Hutchings (eds). Springer Tokyo.  p169-185.

Artois, M., Bengis, R., Delahay, R. J., Duchêne, M. J., Duff, J. P., Ferroglio, E., Gortazar, C., Hutchings, M. R., Kock, R. A., Leighton, F. A., Mörner, T. & Smith, G. C.  (2009).  Wildlife disease surveillance and monitoring.  In Management of Disease in Wild Mammals. R. J. Delahay, G. C. Smith & M. R. Hutchings (eds). Springer Tokyo.  p187-213.

Breed, A. C., Plowright, R. K., Hayman, D. T. S., Knobel, D. L., Molenaar, F. M., Gardner-Roberts, D., Cleaveland, S., Haydon, D. T., Kock, R. A., Cunningham, A. A., Sainsbury, A. W. & Delahay, R. J.  (2009).  Disease management in endangered mammals.  In Management of Disease in Wild Mammals. R. J. Delahay, G. C. Smith & M. R. Hutchings (eds). Springer Tokyo.  p215-239.

Gavier-Widen, D., Chambers, M., Gortazar, C., Delahay, R., Cromie, R. & Linden, A. (2012). Mycobacteria Infections.  In Infectious Diseases of Wild Mammals and Birds in Europe.  D. Gavier-Widen, P. Duff & A. Meredith (eds). Blackwell Publishing Ltd, Oxford. p265-274.

Chambers, M. A., Gormley, E., Corner, L. A. L., Smith, G. C. & Delahay, R. J. (2015). Tuberculosis in Badgers (Meles meles).  In Tuberculosis, Leprosy and Mycobacterial Diseases of Man and Animals: The Many Hosts of Mycobacteria.  H. Mukundan, M. A. Chambers, W. R. Waters & M. Larsen (eds).  CAB International. p296-312.

Palmer, M. V., O’Brien, D. J., Griffin, J. F., Nugent, G., de Lisle, G. W., Ward, A. & Delahay, R. J. (2015). Tuberculosis in Wild and Captive Deer.  In Tuberculosis, Leprosy and Mycobacterial Diseases of Man and Animals: The Many Hosts of Mycobacteria.  H. Mukundan, M. A. Chambers, W. R. Waters & M. Larsen (eds).  CAB International. p334-364..

McDonald, J. L., Delahay, R. J. & McDonald, R. A. (2019).  Bovine tuberculosis in badgers: Sociality, infection and demography in a social mammal. In Wildlife Disease Ecology: Linking Theory to Data and Application.  K. Wilson, A. Fenton & D. Tompkins (eds).  Cambridge University Press.

Scientific papers

Delahay, R.J., Speakman, J.R. & Moss, R. (1995) The energetic consequences of parasitism: effects of a developing infection of Trichostrongylus tenuis (Nematoda) on red grouse (Lagopus lagopus scoticus) energy balance, body weight and condition. Parasitology, 110, 473-482.

Delahay, R. J. & Moss, R. (1996). Food intake, weight changes and egg production in captive red grouse (Lagopus lagopus scoticus) before and during laying: effects of the parasitic nematode Trichostrongylus tenuis. The Condor, 98, 501-511.

Delahay, R. J., Daniels, M. D., Macdonald, D. W., McGuire, K. & D. Balharry. (1998). Do patterns of helminth parasitism differ between groups of wild living cats in Scotland? Journal of Zoology 245 (2), 175-184.

Delahay, R. J., Rogers, L. M., Cheeseman, C. L., Mallinson, P. J., Smith, G. C. and Clifton-Hadley, R. S. (1998).  The transmission of Bovine Tuberculosis between badgers (Meles meles) and domestic cattle in England. Gibier Faune Sauvage, Game Wildl. Vol. 15 (Hors serie Tome 3), p. 805-814.

Delahay, R. J., Cheeseman, C. L., Mallinson, P. J., Rogers, L. M. & Smith, G. C. (1998) Badgers and bovine tuberculosis: A review of studies in the ecology of a wildlife disease reservoir.  Cattle Practice. 6 (2), 83-87.

Rogers, L. M., Delahay, R. J., Cheeseman, C. L.,. Langton, S., Smith, G. C. & Clifton-Hadley, R. (1998) Movement of badgers (Meles meles) in a high density population: Individual, population and disease effects. Proc. Roy. Soc. Ser. B., 265, 1269-1276.

Rogers, L. M., Delahay, R. J., Cheeseman, C. L., Smith, G. C. & Clifton-Hadley, R. S. (1999). The increase in badger (Meles meles) density at Woodchester Park, south west England: a review of the implications for disease (Mycobacterium bovis) prevalence. Mammalia 63(2), 183-192.

Delahay, R.J. (1999) Cestodiasis in the red grouse in Scotland. Journal of Wildlife Diseases 35, 2, 250-258.

Tuyttens, F. A. M., Macdonald, D. W., Delahay, R. J., Rogers, L. M., Mallinson, P. J., Donnelly, C. A. & Newman, C. (1999).  Differences in trappability of European badgers Meles meles in three populations in England.  Journal of Applied Ecology 36, 1051-1062.

Delahay, R. J., Brown, J. A., Mallinson, P. J., Spyvee, P. D., Handoll, D., Rogers, L. M. & Cheeseman, C. L. (2000)  The use of marked bait in studies of the territorial organisation of the European badger (Meles meles). Mammal Review 30(2), 73-87.

Tuyttens, F. A. M., Delahay, R. J., Macdonald, D. W., Cheeseman, C. L., Long, B. and Donnelly, C. (2000). Spatial perturbation caused by a badger (Meles meles) culling operation: implications for the function of territoriality and the control of bovine tuberculosis (Mycobacterium bovis). Journal of Animal Ecology 69(5), 815-828.

Wilkinson, D., Smith, G.C., Delahay, R.J., Rogers, L. M., Cheeseman, C.L. & Clifton-Hadley, R. S. (2000). The effects of bovine tuberculosis (Mycobacterium bovis) on mortality in a badger (Meles meles) population in England. Journal of Zoology 250, 389-395.

Delahay, R. J., Langton, S., Smith, G. C., Clifton-Hadley, R. S. & Cheeseman, C. L. (2000).  The spatio-temporal distribution of Mycobacterium bovis (Bovine Tuberculosis) infection in a high density badger (Meles meles) population. Journal of Animal Ecology 69, 1-15.

Delahay, R. J. & Froelich, K. (2000). Absence of antibodies against canine distemper virus infection in free ranging British populations of the Eurasian badger (Meles meles). Journal of Wildlife Diseases 36(3), 576-579.

Delahay, R. J., Cheeseman, C. L. & Clifton-Hadley, R. S. (2001).  Wildlife disease reservoirs: The epidemiology of Mycobacterium bovis infection in the European badger (Meles meles) and other British mammals.  Tuberculosis 81(1&2), 43-50.

Wilson, G. & Delahay, R. J. (2001). A review of methods to estimate the abundance of terrestrial carnivores using field signs and observation. Wildlife Research 28, 151-164.

Forrester, G., Delahay, R. J. & Clifton-Hadley, R. S. (2001).  Screening badgers (Meles meles) for Mycobacterium bovis using multiple applications of an ELISA test.  Veterinary Record 149(6), 169-172.

Artois, M., Delahay, R., Guberti, V. & Cheeseman, C. (2001). Control of infectious diseases of wildlife in Europe. Veterinary Journal 162, 141-152.

Delahay, R. J., de Leeuw, A. N. S., Claridge, M. C., Harris, A., Cheeseman, C. L., Barlow, A. M., Millar, M. & Clifton-Hadley, R. S. (2001). First report of Mycobacterium bovis in a muntjac deer. Veterinary Record 149(3), 95-96.

Delahay, R. J., de Leeuw, A. N. S., Barlow, A. M., Clifton-Hadley, R. S. & Cheeseman, C. L.  (2002).  The status of Mycobacterium bovis infection in British wild mammals: a review.  Veterinary Journal 164, 90-105.

Garnett, B. T., Delahay, R. J. & Roper, T. J. (2002). Use of cattle farm resources by badgers (Meles meles) and risk of bovine tuberculosis (Mycobacterium bovis) transmission to cattle.  Proceedings of the Royal Society (Series B) 269, 1487-1491.

Garnett, B. T., Roper, T. J. & Delahay, R. J. (2003).  Use of cattle troughs by badgers (Meles meles). A potential route for the transmission of bovine tuberculosis (Mycobacterium bovis) to cattle.  Applied Animal Behaviour Science 80, 1-8.

Wilson, G. J., Delahay, R. J., A. N. S. de Leeuw, P. D. Spyvee & D. Handoll. (2003). Quantification of badger (Meles meles) sett activity as a method of predicting badger numbers.  Journal of Zoology 259, 49-56.

Frantz, A. C., Pope, L. C., Burke, T. A., Wilson, G. J., Delahay, R. J. & Roper, T. J.  (2003).  Reliable microsatellite genotyping of the Eurasian badger (Meles meles) from faecal DNA.  Molecular Ecology 12, 1649-1661.

Roper, T. J., Garnett, B. T. & Delahay, R. J.  (2003).  Visits to farm buildings and cattle troughs by badgers (Meles meles): a potential route for transmission of bovine tuberculosis (Mycobacterium bovis) between badgers and cattle.  Cattle Practice 11, 9-12.

Delahay, R. J., Wilson, G. J., Smith, G. C. & Cheeseman, C. L.  (2003).  Vaccinating badgers (Meles meles) against Mycobacterium bovis: The ecological considerations.  Veterinary Journal 166, 43-51.

Wilson, G. J., Frantz, A. C., Pope, L. C., Roper, T. J., Burke, T. A., Cheeseman, C. L. & Delahay, R. J.  (2003).  Estimation of badger (Meles meles) abundance using faecal DNA typing.  Journal of Applied Ecology 40, 658-666.

Delahay, R. J., Waldram, M., Mallinson, P. J., Spyvee, P. D., Handoll, D., de Leeuw, A. N. S. & Cheeseman, C. L.  (2003).  Response to Tuyttens, Macdonald and Roddam.  Journal of Zoology 260, 119.

de Leeuw, A. N., Forrester, G., Spyvee, P. D., Brash, M. & Delahay, R. J. (2004). Experimental comparison of ketamine with a combination of ketamine, butorphanol and medetomidine for general anaesthesia of the Eurasian badger (Meles meles L.).  Veterinary Journal 167, 186-193.

Hounsome, T., O’Mahony, D. & Delahay, R. (2004).  The diet of Little Owls (Athene noctua) in Gloucestershire, England. Bird Study 51, 282-284.

Wilkinson, D., Smith, G. C., Delahay, R. J. & Cheeseman, C. L.  (2004). A model of bovine tuberculosis in the badger Meles meles: an evaluation of different vaccination strategies.  Journal of Applied Ecology  41, 492-501.

Cagnacci, F., Massei, G., Cowan, D. P. & Delahay, R. J. (2005).  Can learned aversion be used to control bait uptake by Eurasian Badgers?  Applied Animal Behaviour Science 92, 159-168.

Carpenter, P. J., Pope, L. C., Greig, C., Dawson, D. A., Rogers, L. M., Erven, K., Wilson, G. J., Delahay, R. J., Cheeseman, C. L. & Burke, T.  (2005).  Mating system of the Eurasian badger, Meles meles, in a high density population.  Molecular Ecology 14, 273-284.

Delahay, R. J., de Leeuw, A. N., Barlow, A. M., Clifton-Hadley, R. S. & Cheeseman, C. L.  (2005).  The status of Mycobacterium bovis infection in British wild mammals: a review.  Cattle Practice 13, 427-440.

Hounsome, T. D., Young, R. P., Davison, J., Yarnell, R. W., Trewby, I. D., Garnett, B. T., Delahay, R. J. & Wilson, G. J.  (2005).  An evaluation of distance sampling to estimate badger (Meles meles) abundance.  Journal of Zoology 266, 81-87.

Hounsome, T. D. & Delahay, R. J. (2005) Birds in the diet of the Eurasian badger (Meles meles).  A review and a meta-analysis.  Mammal Review 35, 199-209.

Garnett, B. T., Delahay, R. J. & Roper, T. J. (2005). Ranging behaviour of European badgers (Meles meles) in relation to bovine tuberculosis (Mycobacterium bovis) infection. Applied Animal Behaviour Science 94, 331-340.

Delahay, R. J., Smith, G. C., Ward, A. I. & Cheeseman, C. L. (2005).  Options for the Management of Bovine Tuberculosis Transmission from Badgers (Meles meles) to cattle: Evidence from a long-term study.  Mammal Study 30, S73-S81.

Delahay, R. J.  (2006).  Badgers and Bovine Tuberculosis:  The ecological complexities of managing a wildlife disease reservoir.  Cattle Practice 14(1), 7-11.

Anwar, M.A., Knaggs, J., Service, K., McLaren, G. W., Riordan, P., Newman, C., Delahay, R. J., Cheeseman, C. L. & Macdonald, D. W., (2006).  The presence of antibodies to Toxoplasma gondii in European badgers.  Journal of Wildlife Diseases 42, 179-181.

Delahay, R. J., Walker, N., Forrester, G. J., Harmsen, B., Riordan, P., Macdonald, D. W., Newman, C. & Cheeseman, C. L.  (2006).  Demographic correlates of bite wounding in Eurasian badgers (Meles meles L.) in stable and perturbed populations.  Animal Behaviour 71, 1047-1055. 

Young, R. P., Davison, J., Trewby, I. D., Wilson, G. J., Delahay, R. J. & Doncaster, C. P.  (2006).  Abundance of hedgehogs (Erinaceus europaeus) in relation to the density and distribution of badgers (Meles meles). Journal of Zoology 269, 349-356.

Frölich, K., Grabitzky, S. E., Walzer, C., Delahay, R. J., Dorrestein, G. M. & Hatt, J. (2006). Zoo and Wildlife Medical Education: a European Perspective. Journal of Veterinary Medical Education, 33, 401-407.

Courtenay, O., Reilly, L. A., Sweeney, F. P., Hibberd, V., Bryan, S., Ul-Hassan, A., Newman, C., Macdonald, D. W., Delahay, R. J., Wilson, G. J. & Wellington, E. M. H.  (2006).  Is Mycobacterium bovis in the environment important for the persistence of bovine tuberculosis?  Biology Letters 2, 460-462.

Ward, A. I., Tolhurst, B. A. & Delahay, R. J. (2006).  Farm husbandry and the risks of disease transmission between wild and domestic mammals: A brief review focusing on bovine tuberculosis in badgers and cattle.  Animal Science 82: 767-773.

Delahay, R. J., Carter, S.P., Forrester, G. J., Mitchell, A. & Cheeseman, C. L. (2006).  Habitat correlates of group size, bodyweight and reproductive performance in a high-density Eurasian badger (Meles meles) population.  Journal of Zoology 270, 437-447.

Woodroffe, R., Donnelly, C.A., Cox, D.R., Bourne, J.F., Cheeseman, C.L., Delahay, R.J., Gettinby, G., McInerney, J.P. & Morrison, W.I. (2006). Effects of culling on badger Meles meles spatial organization: implications for the control of bovine tuberculosis. Journal of Applied Ecology 43, 1-10.

Woodroffe, R., Donnelly, C. A., Jenkins, H. E., Johnston, W. T., Cox, D. R., Bourne, F. J., Cheeseman, C. L., Delahay, R. J., Clifton-Hadley, R. S., Gettinby, G., Gilks, P., Hewinson, R. G., McInerney, J. P. & Morrison, W. I. (2006).  Culling and cattle controls influence tuberculosis risk for badgers.  Proceedings of the National Academy of Sciences of the United States of America 103, 14713-14717.

Delahay, R. J., Smith, G. C., Barlow, A. M., Walker, N., Harris, A., Clifton-Hadley, R. S. & Cheeseman, C. L.  (2007).  Bovine tuberculosis infection in wild mammals in South West England: A survey of prevalence and a semi-quantitative assessment of the relative risks to cattle. Veterinary Journal 173(2), 287-301.

Delahay, R. J., Ward, A. I., Walker, N., Long, B. & Cheeseman, C. L.  (2007).  The distribution of badger (meles meles) latrines in a high density population: habitat selection and implications for the transmission of bovine tb to cattle.  Journal of Zoology 272, 311-320.

Vicente, J., Delahay, R. J., Walker, N. & Cheeseman, C. L. (2007).  Social organization and movement influence the incidence of bovine tuberculosis in an undisturbed high density badger Meles meles population.  Journal of Animal Ecology 76, 348-360.

Carter, S. P., Delahay, R. J., Smith, G. C., Macdonald, D. W., Riordan, P., Etherington, T. R., Pimley, E. R., Walker, N. J. & Cheeseman, C. L. (2007).  Culling-induced social perturbation in Eurasian badgers Meles meles and the management of TB in cattle: an analysis of a critical problem in applied ecology.  Proceedings of the Royal Society (Series B) 274, 2769-2777.

Cagnacci, F., Massei, G., Cowan, D. P., Walker, N. & Delahay, R. J. (2007).  Effects of bait type and deployment strategy on uptake by free-living badgers. Wildlife Research 34, 454-460.

Courtenay, O., Reilly, L. A., Sweeney, F. P., Macdonald, D. W.,  Delahay, R. J., Wilson, G. J.,  Cheeseman, C. L., Keeling, M. J. & Wellington, E. M. H. (2007).  Performance of an environmental test to detect Mycobacterium bovis infection in badger social groups.  Veterinary Record 161, 817-818.

Pope, L., Butlin, R., Wilson, G., Woodroffe, R., Erven, K., Conyers, C., Franklin, T., Delahay, R., Cheeseman, C. & Burke, T. (2007).  Genetic evidence that culling increases badger movement: implications for the spread of bovine TB.  Molecular Ecology 16, 4919-4929.

Johnson, L. K., Liebana, E., Nunez, A., Spencer, Y., Clifton-Hadley, R., Jahans, K., Ward, A,  Barlow, A. & Delahay, R.  (2008)  Histological observations of bovine tuberculosis in lung and lymph node tissues from British deer. Veterinary Journal 175(3), 409-412.

McDonald, R. A., Delahay, R. J., Carter, S. C., Smith, G. C. & Cheeseman, C. L.  (2008).  Perturbing implications of wildlife ecology for disease control.  Trends in Ecology and Evolution 892, 1-4.

Chambers, M. A., Crawshaw, T., Waterhouse, S., Delahay, R. J., Hewinson, R. G. & Lyashchenko, K. P. (2008).  Validation of the BrockTB Stat-Pak assay for detection of tuberculosis in Eurasian badgers (Meles meles) and influence of disease severity on diagnostic accuracy.  Journal of Clinical Microbiology 46(4), 1498-1500.

Trewby, I., Wilson, G. J., Delahay, R. J., Walker, N., Young,  R., Davison, J., Cheeseman, C., Robertson, P., Gorman, M. & McDonald, R. A.  (2008).  Experimental evidence of competitive release in sympatric carnivores.  Biology Letters 4, 170-172.

Davison, J., Huck, M., Delahay, R. J. & Roper, T. J.  (2008).  Urban badger setts: characteristics, patterns of use and management implications.  Journal of Zoology 275, 190-200.

Winnacker, H., Walker, N., Brash, M. G. I., MacDonald, J. A. & Delahay, R. J. (2008)  Haematological and blood biochemistry values for a wild Eurasian badger (Meles meles) population.  Veterinary Record 162, 551-555.

Tolhurst, B.A; Ward, A.I.; Delahay, R.J.; MacMaster, A. & Roper, T.J. (2008): The behavioural responses of badgers (Meles meles) to exclusion from farm buildings using an electric fence. Applied Animal Behaviour Science 113, 224-235.

Ward, A. I., Tolhurst, B. A., Roper, T. J. & Delahay, R. J. (2008).  Survey of badger access to farm buildings and facilities in relation to contact with cattle.  Vet. Record. 163, 107-111.

Ward, A. I., Pietravalle, S., Cowan, D. P. & Delahay, R. J.  (2008).  Deterrent or dinner bell? Alteration of badger activity and feeding at baited plots using ultrasonic and water jet devices. Applied Animal Behaviour Science 115, 221-232.

Lyashchenko, K. P., Greenwald, R., Esfandiari, J., Chambers, M. A., Vicente, J., Gortazar, C., Santos, N., Correia-Neves, M., Buddle, B. M., Jackson, R., O’Brien, D. J., Schmitt, S., Palmer, M. V., Delahay, R. J. & Waters, W. R. (2008). Animal-side serologic assay for rapid detection of Mycobacterium bovis infection in multiple species of free-ranging wildlife. Veterinary Microbiology 132, 283-292.

Davison, J., Huck, M., Delahay, R. J. & Roper, T. J. (2009). Restricted ranging behaviour in a high density population of urban badgers. Journal of Zoology 277, 45-53.

Delahay, R. J., Davison, J., Poole, D. W, Matthews, A. J., Wilson, C. J., Heydon, M. J. & Roper, T. J. (2009).  Managing conflict between badgers (Meles meles) and humans: recent trends in licensed operations to resolve badger-related problems in England.  Mammal Review 39, 53-66.

Tolhurst, B. A., Delahay, R. J., Walker, N. J., Ward, A. I. & Roper, T. J. (2009). Behaviour of badgers (Meles meles) in farm buildings: opportunities for the transmission of Mycobacterium bovis to cattle? Applied Animal Behaviour Science 117, 103-113.

Stradiotto, A., Cagnacci, F., Delahay, R., Tioli, S., Nieder, L. & Rizzoli, A. (2009).  Spatial organization of the yellow-necked mouse:  Effects of density and resource availability. J. Mammalogy. 90: 704-714.

Ward, A. I., Smith, G. C., Etherington, T. R. & Delahay, R. J. (2009). Estimating the risk of cattle exposure to TB posed by wild deer relative to badgers in England and Wales. Journal of Wildlife Diseases 45 (4), 1104-1120.

Chambers, M. A., Waterhouse, S., Lyashchenko, K., Delahay, R., Sayers, R. & Hewinson, R. G.  (2009).  Performance of TB immunodiagnostic tests in Eurasian badgers (Meles meles) of different ages and the influence of duration of infection on serological sensitivity. BMC Veterinary Research 5: 42. doi:10.1186/1746-6148-5-42.

Ward, A. I., Judge, J. & Delahay, R. J. (2010). Farm husbandry and badger behaviour: opportunities to manage badger to cattle transmission of Mycobacterium bovis? Preventive Veterinary Medicine. 93, 2-10.

Foyle, K. L., Delahay, R. J. & Massei, G. (2010). Isolation of Mycobacterium bovis from a feral wild boar (Sus scrofa) in the UK. Veterinary Record 166, 663-664.

Drewe, J. A., Tomlinson, A., Walker, N. J. & Delahay, R. J. (2010).  Diagnostic accuracy and optimal use of three tests for tuberculosis in live badgers.  PLoS ONE 5(6): e11196. doi:10.1371/journal.pone.0011196

Chambers, M. A., Rogers, F., Delahay, R. J., Lesellier, S., Ashford, R., Dalley, D., Gowtage, S., Davé, D., Palmer, S., Brewer, J., Crawshaw, T., Clifton-Hadley, R., Carter, S., Cheeseman, C., Hanks, C., Murray, A., Palphramand, K., Pietravalle, S., Smith, G. C., Tomlinson, A., Walker, N. J., Wilson, G. J., Corner, L. A. L., Rushton, S. P., Shirley, M. D. F., Gettinby, G., McDonald, R. A. & Hewinson, R. G. (2010).  Bacillus Calmette-Guérin vaccination reduces the severity and progression of tuberculosis in badgers. Proc. R. Soc. B.  278, 1913-1920.  doi:10.1098/rspb.2010.1953

Balseiro, A., Merediz, I., Sevilla, I. A., García-Castro, C., Gortazar, C., Prieto, J. M. & Delahay, R. J.  (2011).  Infection of Eurasian badgers (Meles meles) with Mycobacterium avium complex (MAC) bacteria.  The Veterinary Journal 188, 231-233.  doi: 10.1016/j.tvjl.2010.05.003.

Wilson, G. J., Carter, S. P. & Delahay, R. J.  (2011).  Advances and prospects for management of TB transmission between badgers and cattle.  Veterinary Microbiology 151, 43-50.

Davison, J., Roper, T. J., Wilson, C. J., Heydon, M. J. & Delahay, R. J. (2011).  Assessing spatio-temporal associations in the occurrence of badger-human conflict in England.  European Journal of Wildlife Research. 57(1), 67-76.

Delahay, R. J., Walker, N., Gunn, M. R., Christie, C., Wilson, G. J., Cheeseman, C. L. & McDonald, R. A. (2011).  Using lifetime tooth wear scores to predict age in Eurasian badgers (Meles meles): performance of a predictive model.  Journal of Zoology 796, 1-9.

Palphramand, K. L., Walker, N., McDonald, R. A. & Delahay, R. J. (2011).  Evaluating seasonal bait delivery to badgers using rhodamine B.  European Journal of Wildlife Research 57(1), 35-43.

Tolhurst, B. A., Ward, A. I. & Delahay, R. J. (2011). A study of fox (Vulpes vulpes) visits to farm buildings in Southwest England and the implications for disease management.  European Journal of Wildlife Research 57(6), 1227-1230.

Balseiro, A., Rodríguez, O., González-Quirós, P., Merediz, I., Sevilla, I.A., Davé, D., Dalley, D., Lesellier, S., Chambers, M.A., Bezos, J., Muñoz, M., Delahay, R.J., Gortázar, C. & Prieto, J.M. (2011). Infection of Eurasian badgers (Meles meles) with Mycobacterium bovis and Mycobacterium avium complex in Spain. The Veterinary Journal. doi: 10.1016/j.tvjl.2011.04.012

Travis, E. R., Gaze, W. H., Pontiroli, A., Sweeney, F. P., Porter, D., Mason, S, Keeling, M. J. C., Jones, R. M., Sawyer, J., Aranaz, A., Castellanos Rizaldos, E., Cork, J., Delahay, R. J., Wilson, G. J., Hewinson, R. G., Courtenay, O., & Wellington, E. M. H. (2011).  An Inter-laboratory Validation of a Real Time PCR assay to measure host excretion of bacterial pathogens. PLoS ONE 6(11): e27369. doi:10.1371/journal.pone.0027369

Riordan, P., Delahay, R.J., Cheeseman, C.L., Johnson, P.J., & Macdonald, D.W. (2011).  Culling-induced changes in badger (Meles meles) behaviour, social organisation and the epidemiology of bovine tuberculosis. PLoS ONE 6(12): e28904. doi:10.1371/journal.pone. 0028904

Judge, J., McDonald, R., Walker, N. & Delahay, R. J. (2011).  Effectiveness of exclusion measures in preventing badger visits to farm buildings. PLoS ONE 6(12): e28941. doi:10.1371/journal.pone.0028941

Gortázar, C., Delahay, R., McDonald, R., Boadella, M., Wilson, G., Gavier-Widen, D. & Acevedo, P. (2012). The status of tuberculosis in European mammals.  Mammal Review 42, 193-206.

Mill, A. C., Rushton, S. P., Shirley, M. D. F., Murray, A. W. A., Smith, G. C, Delahay, R. J. & McDonald, R. A. (2012). Farm-scale risk factors for bovine tuberculosis incidence in cattle herds during the Randomised Badger Culling Trial. Epidemiology & Infection 140, 219-230. doi:10.1017/S0950268811000434

Drewe, J. A., Weber, N., Carter, S. P., Bearhop, S., Harrison, X. A., Dall, S. R. X., McDonald, R. A. & Delahay, R. J. (2012).  Performance of proximity loggers in recording intra- and inter-species interactions: a laboratory and field-based validation study.  PLoS ONE. 7(6): e39068.

Palphramand, K., Carter, S., Woods, J., Smith, G., Walker, N. & Delahay, R. (2012).  Uptake of Buried Baits by Badgers: Implications for Rabies Control in Great Britain and the Delivery of an Oral TB Vaccine.  Wildlife Society Bulletin 36, 220-225.  doi: 10.1002/wsb.143.

Tomlinson, A., Chambers, M. & Delahay, R. (2012). Mycobacterium bovis infection in badger cubs: Re-assessing the evidence for maternally derived immunological protection from advanced disease.  Veterinary Immunology and Immunopathology 148, 326-330.

Jenkins, H. E., Cox, D. R. & Delahay, R. J. (2012).  Direction of association between bite wounds and Mycobacterium bovis infection in badgers: implications for transmission. PLoS ONE 7(9), e45584. doi:10.1371/journal.pone.0045584.

Delahay, R. J. (2012).  Guest editorial: A forensic investigation of Mycobacterium bovis infection in badgers. The Veterinary Journal 194, 135-136. http://dx.doi.org/10.101016/j.tvj.2012.08.006.

Carter, S.P., Chambers, M.A., Rushton, S.P., Shirley, M.D.F., Schuchert, P., Pietravalle, S., Murray, A., Rogers, F., Gettinby, G., Smith, G.C., Delahay, R.J., Hewinson, R.G. & McDonald, R.A. (2012) BCG vaccination reduces risk of tuberculosis infection in vaccinated badgers and unvaccinated badger cubs. PLOS ONE. 7(12): e49833. doi:10.1371/journal.pone.0049833

Weber, N., Bearhop, S., Dall, S. R. X., Delahay, R. J., McDonald, R. A. & Carter, S. P. (2012).  Denning behaviour in the European badger (Meles meles) correlates with bovine tuberculosis infection status. Behavioural Ecology & Sociobiology 67, 471-479.  doi: 10.1007/s00265-012-1467-4

Newth, J. L., Cromie, R. L., Brown, M. J., Delahay, R. J., Meharg, A. A., Deacon, C., Norton, G. J., O’Brien, M. F. & Pain, D. J. (2013).  Poisoning from lead gunshot: still a threat to wild waterbirds in Britain.  European Journal of Wildlife Research 59,195–204. doi: 10.1007/s10344-012-0666-7.

Robertson, A., McDonald, R. A., Delahay, R. J., Kelly, S. D. & Bearhop, S.  (2013).  Whisker growth in wild Eurasian badgers Meles meles: implications for stable isotope and bait-marking studies.  European Journal of Wildlife Research 59, 341-350.

Graham, J., Smith, G., Delahay, R. J., Bailey, T.C., McDonald, R. & Hodgson, D. (2013).  Multi-state modelling reveals sex-dependent transmission progression and severity of tuberculosis in wild badgers. Epidemiology and Infection 141, 1429-1436.  doi:10.1017/S0950268812003019.

Balseiro, A., González-Quirós, P., Rodríguez, O., Francisca Copano, M., Merediz, I., de Juan, L., Chambers, M. A., Delahay, R. J., Marreros, N., Royo, L. J., Gortázar, C. & Prieto, J. M. (2013).  Spatial relationships between Eurasian badgers (Meles meles) and cattle infected with Mycobacterium bovis in Northern Spain.  Veterinary Microbiology 197(3), 739-745.  doi: 10.1016/j.tvjl.2013.03.017.

Mardare, C., Delahay, R. J. & Dale, J. W. (2013). Environmental amoebae do not support the long-term survival of virulent mycobacteria.  Journal of Applied Microbiology 114(5), 1388-1394.  doi: 10.1111/jam.12166.

Jones, R. M., Ashford, R., Cork, J., Palmer, S., Wood, E., Spyvee, P., Parks, S., Bennett, A., Brewer, J., Delahay, R., Chambers M. & Sawyer, J. (2013).  Evaluation of a method to detect Mycobacterium bovis in air samples taken from infected badgers (Meles meles) and their setts.  Letters in Applied Microbiology 56, 361-365.

Tomlinson, A. J., Chambers, M. A., Carter, S. P., Wilson, G., Smith, G., McDonald, R. A., & Delahay, R. J.  (2013).   Heterogeneity in the risk of Mycobacterium bovis infection in European badger (Meles meles) cubs.  Epidemiology & Infection 141, 1458-1466.  doi: 10.1017/S095026881300040X.

Drewe, J. A., O’Connor, H. M., Weber, N., McDonald, R. A. & Delahay, R. J. (2013). Patterns of direct and indirect contact among cattle and badgers naturally infected with tuberculosis: heterogeneity in individual behaviour and tuberculosis transmission risks.  Epidemiology & Infection 141, 1467-1475.  doi: 10.1017/S0950268813000691.

Delahay, R. J., Walker, N., Smith, G. S., Wilkinson, D., Clifton-Hadley, R. S., Cheeseman, C. L., Tomlinson, A. J. & Chambers, M. A. (2013).  Long-term temporal trends and estimated transmission rates for Mycobacterium bovis infection in an undisturbed high density badger (Meles meles) population.  Epidemiology & Infection 141, 1445–1456. doi:10.1017/S0950268813000721

Weber, N., Carter, S. P., Dall, S. R. X., Delahay, R. J., McDonald, J. L., Bearhop, S. & McDonald, R. A. (2013).  Badger social networks correlate with tuberculosis infection.  Current Biology 23(20), R1-R2.

Tomlinson, A.J., Chambers, M.A., Wilson, G.J., McDonald, R.A. and Delahay, R.J.  (2013).  Sex-related heterogeneity in the life-history correlates of Mycobacterium bovis infection in European badgers (Meles meles).  Transboundary and Emerging Diseases 60 (Suppl. 1), 37-45.

Judge, J., Wilson, G. J., Macarthur, R., Delahay, R. J. & McDonald, R. A.  (2013).  Density and abundance of badger social groups in England and Wales in 2011-2013.  Scientific Reports 4, 3809. doi: 10.1038/srep03809.

Robertson, A., McDonald, R. A., Delahay, R. J., Kelly, S. & Bearhop, S. (2014).  Individual foraging specialisation in a social mammal: the European badger (Meles meles).  Oecologia doi 10.1007/s00442-014-3019-2.

Trewby, I. D., Young, R., McDonald, R. A., Wilson, G. J., Davison, J., Walker, N., Robertson, A., Doncaster, P. & Delahay, R. J.  (2014).  Impacts of removing badgers on localised counts of hedgehogs.  PLoS ONE 9 (4), e95477. Doi: 10.1371/journal.pone.0095477.

Gortazar, C., Reperant, L.A., Kuiken, T., de la Fuente, J., Boadella, M., Martínez, B., Ruiz-Fons, J.F., Estrada-Peña, A., Drosten, C., Medley, G., Ostfeld, R., Peterson, T., VerCauteren, K., Menge, C., Artois, M., Schultsz, C., Delahay, R., Serra-Cobo, J., Poulin, R., Keck, F., Aguirre, A.A., Henttonen, H., Dobson, A.P., Kutz, S., & Lubroth, J.  (2014).  Crossing the Interspecies Barrier: Opening the Door to Zoonotic Pathogens. PLoS Pathogens 10(6), e1004129.

McDonald, J. L., Smith, G. C., McDonald, R. A., Delahay, R. J. & Hodgson, D. (2014).  Mortality trajectory analysis reveals the drivers of sex-specific epidemiology in natural wildlife-disease interactions. Proceedings of the Royal Society Series B.281:20140526.

Tomlinson, A. J., Chambers, M. A., McDonald, R. A. & Delahay, R. J.  (2014).  Association of quantitative interferon-c responses with the progression of naturally acquired Mycobacterium bovis infection in wild European badgers (Meles meles).  Immunology 144, 263–270.

Robertson, A. B., Palphramand, K. L., Carter, S. P. & Delahay, R. J. (2014).  Group size correlates with territory size in European badgers: implications for the resource dispersion hypothesis?  Oikos doi: 10.1111/oik.01459.

Beirne, C., Delahay, R., Hares, M. & Young, A. (2014).  Age-related declines and disease-associated variation in immune cell telomere length in a wild mammal. PLoS ONE 9(9): e108964. doi:10.1371/journal.pone.0108964.

Benton, C., Delahay, R., Trewby, H. & Hodgson, D. (2014).  What has molecular epidemiology ever done for wildlife disease research?  Past contributions and future directions.  European Journal of Wildlife Research 61: 1-16.  doi 10.1007/s10344-014-0882-4.

Muñoz-Mendoza, M., Romero, B., del Cerro, A., Gortázar, C., García-Marín, J. F., Menéndez, S., Mourelo, J., de Juan, L., Sáez, J. L., Delahay, R. J. & Balseiro, A.  (2015).  Sheep as a potential source of bovine TB: epidemiology, pathology and evaluation of diagnostic techniques. Transboundary and Emerging Diseases.  doi: 10.1111/tbed.12325.

Robertson, A., Chambers, M. A., Delahay, R. J., McDonald, R. A., Palphramand, K. L., Rogers, F. & Carter, S. P.  (2015).  Exposure of non-target wildlife to candidate TB vaccine baits deployed for European badgers. European Journal of Wildlife Research 61:263–269.  doi 10.1007/s10344-014-0896-y.

Robertson, A., McDonald, R. A., Delahay, R. J., Kelly, S. & Bearhop, S. (2015).  Resource availability affects individual niche variation and its consequences in group-living European Badgers Meles melesOecologia 178: 31-43.  doi 10.1007/s00442-014-3202-5.

McClune, D. W., Marks, N. J., Delahay, R. J., Montgomery, W. I. & Scantlebury, D. M. (2015).  Behaviour-time budget and functional habitat use of a free-ranging European badger (Meles meles).  Animal Biotelemetry 3:7. doi: 10.1186/s40317-015-0025-z

Delahay, R.  (2015). Control of bovine tuberculosis in New Zealand in the face of a wildlife host: a compiled review of 50 years of programme policy, design and research.  New Zealand Veterinary Journal 63:sup1, 2-3, DOI: 10.1080/00480169.2015.1047435

King, H., Murphy, A., James, P., Travis, E., Porter, D., Sawyer, J., Cork, J., Delahay, R. J., Gaze, W., Courtenay, O. & Wellington, E. M. (2015).  Performance of a non-Invasive test for detecting Mycobacterium bovis shedding in European badger (Meles meles) populations.  Journal of Clinical Microbiology 53 (7), 2316-2323. doi: 10.1128/JCM.00762-15.

Beirne, C., Delahay, R. & Young, A. (2015).  Sex differences in senescence: the role of intra-sexual competition in early adulthood.  Proceedings of the Royal Society (Series B) 282: 20151086.  DOI 10.1098/rspb.2015.1086

McClune, D. W. Kostka, B., Delahay, R. J., Montgomery, W. I., Marks, N. J. & Scantlebury, D. M. (2015).  Winter is coming: seasonal variation in resting metabolic rate of the European badger (Meles meles).  PLoS ONE. Published: September 9, 2015: DOI: 10.1371/journal.pone.0135920

King, H., Murphy, A., James, P., Travis, E., Porter, D., Hung, Y., Sawyer, J., Cork, J., Delahay, R. J., Gaze, W., Courtenay, O. & Wellington, E. M. (2015).  The variability and seasonality of the environmental reservoir of Mycobacterium bovis shed by wild European badgers. Scientific Reports. Aug 6;5:12318. doi: 10.1038/srep12318.

Ideozu, E. J., Whiteoak, A. M., Tomlinson, A. J., Robertson, A., Delahay, R. J. & Hide, G. (2015).  High prevalence of trypanosomes in European badgers detected using ITS-PCR. Parasites and Vectors 8:480. DOI 10.1186/s13071-015-1088-7

Buzdugan, S. N., Chambers, M. A., Delahay, R. J. & Drewe, J. A. (2016).  Diagnosis of tuberculosis in groups of badgers: An exploration of the impact of trapping efficiency, infection prevalence and the use of multiple tests.  Epidemiology & Infection available on CJO2016. doi:10.1017/S0950268815003210.

McDonald, J. L., Bailey, T., Delahay, R. J., McDonald, R. A., Smith, G. C. & Hodgson, D. J.  (2016).  Demographic buffering and compensatory recruitment promotes the persistence of disease in a wildlife population.  Ecology Letters. doi: 10.1111/ele.12578.

Beirne, C., Waring, L., McDonald, R. A., Delahay, R. & Young, A.  (2016).  Age-related declines in immune response in a wild mammal are unrelated to immune cell telomere length.  Proceedings of the Royal Society (Series B). 283: doi: 10.1098/rspb.20152949.

Broughan, J. M., Judge, J., Ely, E., Delahay, R. J., Wilson, G., Clifton-Hadley, R. S., Goodchild, A. V., Bishop, H., Parry, J. E. & Downs, S. H.  (2016). A Review of risk factors for bovine tuberculosis infection in cattle in the UK and Ireland. Epidemiology & Infection.  Doi: 10.1017/S095026881600131X.

Benton, C. H., Delahay, R. J., Robertson, A., McDonald, R. A., Wilson, A. J., Burke, T. A. & Hodgson, D.  (2016).  Blood thicker than water: Kinship, disease prevalence and group size drive divergent patterns of infection risk in a social mammal.  Proceedings of the Royal Society (Series B). 283: 20160798;  DOI: 10.1098/rspb.2016.0798. 

Ward, A. I., Finney, J. K., Beatham, S. E., Delahay, R. J., Robertson, P. A. & Cowan, D. P. (2016).  Exclusions for resolving urban badger damage problems: outcomes and consequences.  PeerJ. 4: e2579; DOI 10.7717/peerj.2579.

Robertson, A., Delahay, R., Wilson, G., Vernon, I., Judge, J. & McDonald, R. A. (2017).  How well do farmers know their badgers?  Relating farmer knowledge to ecological survey data.  Veterinary Record 180:48. DOI:10.1136/vr.103819.

Robertson, A., Delahay, R., McDonald, R. A., Aylett, P., Henderson, R. J., Gowtage, S., Chambers, M. A. & Carter, S. P.  (2016).  Behaviour of European badgers and non-target species towards candidate baits for oral delivery of a tuberculosis vaccine.  Preventative Veterinary Medicine 135: 95–101. DOI 10.1016/j.prevetmed.2016.11.007.

Smith, G. C., Delahay, R. J., McDonald, R. A. & Budgey, R.  (2016). Model of selective and non-selective management of badgers (Meles meles) to control bovine tuberculosis in badgers and cattle.  PLoS ONE. 11(11): e0167206. doi:10.1371/journal.pone.0167206.

Silk, M. J., Croft, D. P., Delahay, R. J., Hodgson, D. J., Boots, M. & McDonald, R. A. (2017). Using social network measures in wildlife disease ecology, epidemiology and management.  Bioscience biw175. DOI: https://doi.org/10.1093/biosci/biw175.

Judge, J., Wilson, G. J., Macarthur, R., McDonald, R. A. & Delahay, R. J. (2017).  Abundance of badgers (Meles meles) in England and Wales. Scientific Reports 7: 276. DOI:10.1038/s41598-017-00378-3.

Buzdugan, S. N., Vergne, T., Grosbois, V., Delahay, R. J. & Drewe, J. A. (2017).  Inference of the infection status of individuals using longitudinal testing data from cryptic populations: Towards a probabilistic approach to diagnosis.  Scientific Reports 7: 1111.  DOI:10.1038/s41598-017-00806-4

Silk, M. J., Croft, D. P., Delahay, R. J., Hodgson, D. J., Boots, M. & McDonald, R. A. (2017). The application of statistical network models in disease research.  Methods in Ecology & Evolution. DOI: 10.1111/2041-210X.12770

Stewart, L. D., Tort, N., Meakin, P., Argudo, J. M., Nzuma, R., Reid, N., Delahay, R. J., Ashford, R., Montgomery, W. I. & Grant, I. R. (2017). Development of a novel immunochromatographic lateral flow assay specific for Mycobacterium bovis cells and its application in combination with immunomagnetic separation to test badger faeces. BMC Veterinary Research 13: 131. DOI: 10.1186/s12917-017-1048-x

Palphramand, K., Delahay, R., Robertson, A., Gowtage, S., Williams, G. A., McDonald, R. A., Chambers, M. & Carter, S. P.  (2017). Field evaluation of candidate baits for oral delivery of BCG vaccine to European badgers Meles melesVaccine 35: 4402–4407.  DOI: 10.1016/j.vaccine.2017.06.059

Buzdugan, S. N., Chambers, M. A., Delahay, R. J. & Drewe, J. A. (2017).  Quantitative interferon-gamma responses predict future disease progression in badgers naturally infected with Mycobacterium bovisEpidemiology & Infection 145(15):3204-3213.  DOI 10.1017/S0950268817001960

Silk, M. J., Weber, N., Steward, L. C., Delahay, R. J., Croft, D. P., Hodgson, D. J., Boots, M. & McDonald, R. A.  (2017).  Seasonal variation in daily patterns of social contacts in the European badger Meles melesEcology & Evolution 7(21), 9006–9015.  DOI 10.1002/ece 3.3402

Silk, M., Weber, N. L., Steward, L. C., Hodgson, D., Boots, M., Croft, D. P., Delahay, R. J. & McDonald, R. A. (2018).  Contact networks structured by sex underpin sex-specific epidemiology of infection.  Ecology Letters 21, 309–318. DOI 10.1111/ele.12898

Silk, M. J., Drewe, J. A., Delahay, R. J., Weber, N., Steward, L. C., Wilson-Aggarwal, J., Boots, M., Hodgson, D. J., Croft, D. P. &. McDonald, R. A. (2018).  Quantifying direct and indirect contacts for the potential transmission of infection between species using a multilayer contact network. Behaviour 15: 731-757.  DOI10.1163/1568539X-00003493.

Benton, C., Delahay, R., Smith, F., Robertson, A., McDonald, R., Burke, T. & Hodgson, D.  (2018) Inbreeding intensifies sex- and age-dependent disease in a wild mammal. Journal of Animal Ecology 87, 1500-1511. doi.org/10.1111/1365-2656.12878.

Whiteoak, A. M., Ideozu, E. J., Alkathiry, H., Tomlinson, A. J., Delahay, R. J., Cowen, S., Mullineaux, E., Gormley, E., Birtles, R. J., Lun, Z. R. & Hide, G.  (2018).  Investigation into the genetic diversity in Toll-like Receptors 2 and 4 in the European badger Meles meles. Research in Veterinary Science 119, 228–231.  doi.org/10.1016/j.rvsc.2018.06.020.

Rozins, C., Silk, M., Croft, D. P., Delahay, R. J., Hodgson, D., McDonald, R. A., Weber, N. & Boots, M. (2018).  Social structure contains epidemics and regulates individual roles in disease transmission in a group-living mammal.  Ecology and Evolution 2018; 00:1–12. https://doi.org/10.1002/ece3.4664  

Smith, G. C. & Delahay, R. J. (2018).  Modelling as a decision support tool for bovine TB control programs in wildlife.  Frontiers in Veterinary Science 5: 276. doi: 10.3389/fvets.2018.00276

Colchero, F., Jones, O. R., Conde, D. A., Hodgson, D., Zajitschek, F., Schmidt, B. R., Malo, A. F., Alberts, S. C., Becker, P. H., Bouwhuis, S., Bronikowski, A. M., De Vleeschouwer, K. M., Delahay, R. J., Dummermuth, S., Fernández-Duque, E., Flatt, T., Frisenvænge, J., Hesselsøe, M., Larson, S., Lemaître, J., McDonald, J., Miller, D. A. W.,  O'Donnell, C., Packer, C., Raboy, B. E., Reading, C. J., Wapstra, E., Weimerskirch, H., While, G. M., Baudisch, A., Coulson, T. & Gaillard, J. (2018).  The diversity of population responses to environmental change.  Ecology Letters 22: 342–353. doi: 10.1111/ele.13195

Carter, S. P., Robertson, A., Palphramand, K. L., Chambers, M. A., McDonald, R. A. & Delahay, R. J. (2018).  Bait uptake by wild badgers and its implications for oral vaccination against tuberculosis.  PLoS ONE 13 (11): e0206136. doi: 10.1371/journal. pone.0206136

Marjamäki, P. H., Dugdale, H. L., Dawson, D. A., McDonald, R. A., Delahay, R., Burke, T. & Wilson, A. J. (2019).  Individual variation and the source-sink group dynamics of extra-group paternity in a social mammal.  Behavioral Ecology 30 (2), p301–312. doi.org/10.1093/beheco/ary164

Fielding, H. R., McKinley, T. J., Silk, M. J., Delahay, R. J. & McDonald, R. A. (2019).  Contact chains of cattle farms in Great Britain.  Royal Society Open Science 6: 180719. doi 10.1098/rsos.180719

Silk, M. J., Hodgson, D. J., Rozins, C., Croft, D. P., Delahay, R. J., Boots, M. & McDonald, R. A. (2019).  Integrating social behaviour, demography and disease dynamics in network models: applications to disease management in declining wildlife populations.  Philosophical Transactions. B 374: 20180211. http://dx.doi.org/10.1098/rstb.2018.0211

Robertson, A., Judge, J., Wilson, G. J., Vernon, I., Delahay, R. J. & McDonald, R. A.  (2019).  Predicting badger visits to farm yards and making predictions available to farmers.  PLoS ONE. 14(5):e0216953. doi10.1371/journal.pone.0216953

Chen, R., Little, R., Mihaylova, L., Delahay, R. & Cox, R.  (2019).  Wildlife surveillance using deep learning methods. Ecology and Evolution 9:9453-9466. DOI: 10.1002/ece3.5410

Barbour, K., McClune, D. W., Delahay, R. J., Speakman, J. R., Kostka, B., Montgomery, W. I., Marks, N. J. & Scantlebury, D. M. (2019).  No energetic cost of tuberculosis infection in European badgers (Meles meles).  Journal of Animal Ecology DOI: 10.1111/1365-2656.13092

Hudson, D. W., Delahay, R. J., McDonald, R., McKinley, T. J. & Hodgson, D. J. (in press).  Analysis of lifetime mortality trajectories in wildlife disease research: BaSTA and beyond.  Diversity.

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